1. Institut Jean-Pierre Bourgin; INRA; AgroParisTech; CNRS; Université Paris-Saclay; RD10 78026 Versailles Cedex France
2. Institute for Plant Biotechnology; Department of Genetics; Stellenbosch University; Private Bag X1 Matieland 7602 South Africa
3. School of Biological Sciences; University of Bristol; Life Sciences Building 24 Tyndall Avenue Bristol BS8 1TQ UK
4. Laboratoire de Reproduction et Développement des Plantes; Université de Lyon; Ecole Normale Supérieure de Lyon; Université Claude Bernard Lyon 1; CNRS, INRA; Lyon 69364 France