1. College of AgronomyHunan Agricultural University Changsha Hunan 410128 China
2. National Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan 430070 China
3. National Key Laboratory of Wheat and Maize Crops Science College of Agronomy Henan Agricultural University Zhengzhou 450002 China
4. Key Laboratory of Horticultural Plant Biology (Ministry of Education) Huazhong Agricultural University Wuhan 430070 China