1. Department of Microbiology; Meiji Pharmaceutical University; Kiyose Tokyo 204-8588 Japan
2. Department of Dermatology; Tokyo Medical University; Shinjuku-ku Tokyo 160-0023 Japan
3. Japan Collection of Microorganisms; RIKEN (The Institute of Physical and Chemical Research); Wako Saitama 351-0198 Japan
4. Department of Immunobiology; Meiji Pharmaceutical University; Kiyose Tokyo 204-8588 Japan