1. Department of Biological Science and Technology, Faculty of Engineering; The University of Tokushima; Tokushima Tokushima 770-8506 Japan
2. Department of Clinical and Diagnostic Oral Sciences; Queen Mary's School of Medicine and Dentistry; London E1 2AD United Kingdom
3. MRC Molecular Pathogenesis Group, Barts and The London School; Queen Mary's School of Medicine and Dentistry; London E1 2AD United Kingdom
4. Graduate School of Bioagricultural Sciences; Nagoya University; Nagoya Aichi 464-8601 Japan
5. Department of Microbiology, Institute of Health Biosciences; The University of Tokushima Graduate School; Tokushima Tokushima 770-8504 Japan