1. Departments of Bioregulation, Leprosy Research Center; National Institute of Infectious Diseases; Higashimurayama Tokyo 189-0002 Japan
2. Microbiology, Leprosy Research Center; National Institute of Infectious Diseases; Higashimurayama Tokyo 189-0002 Japan
3. Department of Bacteriology; Yokohama City University School of Medicine; Yokohama Kanagawa 236-0004 Japan