1. Organismal and Evolutionary Biology Research Programme, Faculty of Biological and Environmental Sciences University of Helsinki Helsinki Finland
2. Department of Biology and Environmental Science University of Jyväskylä Jyväskylä Finland
3. College Life and Environmental Sciences, Penryn Campus University of Exeter Penryn UK
4. Department of Interdisciplinary Life Sciences Konrad Lorenz Institute of Ethology, University of Veterinary Medicine Vienna Vienna Austria
5. Institute for Biodiversity and Ecosystem Dynamics University of Amsterdam Amsterdam The Netherlands