1. Graduate School of Life and Environmental Sciences; University of Tsukuba; Ibaraki; Japan
2. Institute of Arid Region; Medenine; Tunisia
3. Alliance for Research on North Africa (ARENA); University of Tsukuba; Ibaraki; Japan
4. Laboratoire des Plantes Extrêmophiles; Centre de Biotechnologie à la Technopole de Borj-Cédria(CBBC); Hammam-lif; Tunisia