1. Department of Ecology and Genetics; Evolutionary Biology Centre; Science for Life Laboratory; Uppsala University; Norbyvägen 18D 75236 Uppsala Sweden
2. Institute of Biosciences and Bioresources; Division of Florence; National Research Council; Via Madonna del Piano 10 50019 Sesto Fiorentino (Firenze) Italy
3. Institute of Plant and Animal Ecology; Ural Branch of Russian Academy of Science; Ekaterinburg 620144 Russia