1. CAS Key Laboratory of Forest Ecology and Management Institute of Applied Ecology, Chinese Academy of Sciences Shenyang China
2. Department of Forest Vegetation Forestry and Forest Products Research Institute Tsukuba Japan
3. CAS Key Laboratory of Tropical Plant Resources and Sustainable Use Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences Menglun China
4. Center for Plant Ecology, Core Botanical Gardens Chinese Academy of Sciences Xishuangbanna China
5. Organismal and Evolutionary Biology, Viikki Plant Science Centre (ViPS) University of Helsinki Helsinki Finland
6. Graduate School of Life Sciences Tohoku University Sendai Japan