1. Laboratory of Ecology and Evolutionary Biology, School of Ecology and Environmental Sciences Yunnan University Kunming 650500 China
2. Institute of International Rivers and Eco‐Security Yunnan University Kunming 650500 China
3. Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Biology Yunnan University Kunming 650500 China
4. Computational Biology and Population Genomics Group, Departamento de Biologia Animal, Faculdade de Ciências, Centre for Ecology, Evolution and Environmental Changes Universidade de Lisboa 1649‐004 Lisbon Portugal
5. School of Food Science Tibet Agricultural and Animal Husbandry University Nyingchi 860000 China
6. Division of Evolutionary Ecology, Institute of Ecology and Evolution University of Bern 3012 Bern Switzerland