1. The Key Laboratory of Plant Development and Environmental Adaptation Biology Ministry of Education School of Life Science Shandong University (Qingdao) Qingdao 266237 China
2. Department of Molecular Nutrition and Biochemistry of Plants Institute of Plant Nutrition Leibniz University Hannover Hannover 30419 Germany
3. Shanghai Center for Plant Stress Biology & National Key Laboratory of Plant Molecular Genetics Center for Excellence in Molecular Plant Sciences Chinese Academy of Sciences Shanghai 200032 China
4. School of Life Science Shandong University (Qingdao) Qingdao 266237 China
5. School of Environmental Science and Engineering Shandong University (Qingdao) Qingdao 266237 China
6. Key Laboratory of Vegetation Ecology Ministry of Education Jilin Songnen Grassland Ecosystem National Observation and Research Station Institute of Grassland Science Northeast Normal University Changchun 130024 China