1. University of Jyvaskyla; Department of Biological and Environmental Science; P.O. Box 35 FI-40014 University of Jyvaskyla Finland
2. Department of Biology; University of Turku; Turku Finland
3. University of Jyvaskyla; Centre of Excellence in Biological Interactions; P.O. Box 35 FI-40014 University of Jyvaskyla Finland