1. Ecological Genetics Laboratory; Center for Frontier Research; National Institute of Genetics; Mishima Japan
2. Atmosphere and Ocean Research Institute; The University of Tokyo; Kashiwanoha Japan
3. Biological Laboratory; Gifu-Keizai University; Ogaki Japan
4. Department of Applied Aquabiology; National Fisheries University; Shimonoseki Japan
5. PRESTO; Japan Science and Technology Agency; Honcho Kawaguchi Japan