1. School of Biological Sciences; University of Sydney; Sydney NSW 2006 Australia
2. Smithsonian Tropical Research Institute; Apartado 0843-03092 Balboa Ancon Republic of Panama
3. Berrimah Veterinary Laboratories; Northern Territory, Department of Primary Industries and Fisheries; Berrimah NT 0828 Australia
4. School of Biosciences; University of Melbourne; Melbourne VIC 3010 Australia