1. CSIRO Land & Water; Tropical Ecosystems Research Centre; PMB 44 Winnellie NT 0822 Australia
2. Center for Macroecology, Evolution, and Climate; Department of Biology; University of Copenhagen; Universitetsparken 15 Copenhagen DK-2100 Denmark
3. School of Environmental Sciences; University of Liverpool; Liverpool L69 3GP UK
4. School of Animal; Plant and Environmental Sciences; University of the Witwatersrand; Private Bag X3 Wits 2050 South Africa