1. Southern Marine Science and Engineering Guangdong Laboratory Zhanjiang China
2. Southern Marine Science and Engineering Guangdong Laboratory Zhuhai China
3. Institute of Modern Aquaculture Science and Engineering School of Life Sciences South China Normal University Guangzhou China
4. Key Laboratory of Eutrophication and Red Tide Prevention of Guangzhou Higher Education Institutes Research Center of Harmful Algae and Marine Biology Jian University Guangzhou China