1. Department of Biological Sciences University of Idaho Moscow ID 83844 USA
2. Institute for Bioinformatics and Evolutionary Studies University of Idaho Moscow ID 83844 USA
3. Florida Museum of Natural History University of Florida Gainesville FL 32611 USA
4. Department of Biology University of Florida Gainesville FL 32611 USA
5. Department of Ecology and Evolutionary Biology University of Michigan Ann Arbor MI 48109 USA