1. College of Life Sciences; Hunan Normal University; Changsha China
2. Centre for Behavioural Ecology and Evolution; College of Life Sciences; Hubei University; Wuhan China
3. Department of Zoology; National Museum of Nature and Science; 4-1-1 Amakubo Tsukuba-shi Ibaraki-ken 305-0005 Japan
4. Department of Entomology; National Museum of Natural History, Smithsonian Institution; Washington D.C. USA
5. Department of Biology; University of Vermont; 109 Carrigan Drive MLS-316 Burlington VT 05405 USA
6. Department of Biological Sciences; National University of Singapore; Singapore
7. Evolutionary Zoology Laboratory; Biological Institute ZRC SAZU; Ljubljana Slovenia