1. Wildlife Ecology; Department of Bioscience; Aarhus University; Kalø, Grenåvej 14 DK-8410 Rønde Denmark
2. Ecoinformatics and Biodiversity; Department of Bioscience; Aarhus University; Ny Munkegade 114 DK-8000 Aarhus C Denmark
3. School Biology and Ecology and Sustainability Solutions Initiative; University of Maine; Deering 303 Orono ME 04469 USA