1. Department of Earth and Environmental Sciences; University of Milano Bicocca; Piazza della Scienza 1 20126 Milano Italy
2. Laboratoire d'Ecologie Alpine (LECA); Université Grenoble Alpes; F-38000 Grenoble France
3. CNRS; LECA; F-38000 Grenoble France
4. Global Mammal Assessment Program; Department of Biology and Biotechnologies; Sapienza University of Rome; Viale dell'Università 32 I-00185 Roma Italy