1. Centre d'Ecologie Fonctionnelle et Evolutive CEFE‐CNRS Université Montpellier UMR 5175 Montpellier 34293 France
2. Department of Ecology and Evolutionary Biology Princeton University Princeton NJ 08544 USA
3. Percy FitzPatrick Institute DST/NRF Centre of Excellence University of Cape Town Rondebosch 7701 South Africa