1. Institut de Biologie Intégrative et des Systèmes, Pavillon Charles-Eugène-Marchand, 1030 Avenue de la Médecine; Université Laval; Québec Québec G1V 0A6 Canada
2. Present Address: Coastal Oregon Marine Experiment Station, Hatfield Marine Science Center; Oregon State University; 2030 SE Marine Science Dr Newport Oregon 97365
3. Department of Arctic and Marine Biology; University of Tromsø; N9037 Tromsø Norway