1. Key Laboratory of Bio-Resources and Eco-Environment of Ministry of Education, College of Life Sciences; Sichuan University; Chengdu 610064 China
2. Floricultural Research Institute; Guangdong Academy of Agricultural Sciences; Guangzhou Guangdong 510640 China
3. Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden; the Chinese Academy of Sciences; Mengla 666303 China
4. Graduate School of the Chinese Academy of Sciences; Beijing 100039 China