1. School of Life Sciences University of Nottingham Nottingham UK
2. Centre for Sport, Exercise and Osteoarthritis Research Versus Arthritis Nottingham UK
3. MRC Versus Arthritis Centre for Musculoskeletal Ageing Research Nottingham UK
4. School of Medicine University of Nottingham Nottingham UK
5. NIHR Nottingham Biomedical Research Centre Nottingham UK