1. Southern Regional Collaborative Innovation Center for Grain and Oil Crops in China College of Agriculture Hunan Agricultural University Changsha 410128 China
2. Department of Biology Hong Kong Baptist University Kowloon, Hong Kong China
3. School of Life Sciences and State Key Laboratory of Agrobiotechnology The Chinese University of Hong Kong Shatin, Hong Kong China
4. Key Laboratory of Crop Genetics and Physiology of Jiangsu Province Yangzhou University Yangzhou 225000 China
5. College of Bioscience and Biotechnology Hunan Agricultural University Changsha 410128 China