1. Ministry of Education Key Laboratory of Cell Activities and Stress Adaptations; School of Life Sciences; Lanzhou University; Lanzhou 730000 China
2. College of Agronomy; Sichuan Agricultural University; Chengdu 611130 China
3. College of Life Sciences; Inner Mongolia Agricultural University; Hohhot 010018 China
4. Ministry of Education Key Laboratory of Bio-Resource and Eco-Environment; College of Life Sciences; Sichuan University; Chengdu 610064 China