1. School of Biological Sciences University of Aberdeen Aberdeen AB24 3UU UK
2. The James Hutton Institute Craigiebuckler Aberdeen AB15 8QH UK
3. School of Geosciences University of Edinburgh Edinburgh EH9 3FF UK
4. Department of Earth and Environmental Sciences University of Manchester Manchester M13 9PT UK
5. Scottish Natural Heritage The Links Golspie KW10 6UB UK