1. State Key Laboratory of Plant Cell and Chromosome Engineering; Institute of Genetics and Developmental Biology; Chinese Academy of Sciences; Beijing 100101 China
2. University of Chinese Academy of Sciences; Beijing 100039 China
3. Shanxi Agriculture University; Taigu 030801 China
4. State Key Laboratory for Biology of Plant Diseases and Insect Pest; Institute of Plant Protection; Chinese Academy of Agricultural Sciences; Beijing 100193 China
5. Institute for Plant Breeding, Genetics and Genomics; University of Georgia; 111 Riverbend Road Athens GA 30602 USA