1. School of Environmental Sciences; University of Liverpool; Liverpool L69 3GP UK
2. NERC Centre for Ecology and Hydrology; Maclean Building Benson Lane Crowmarsh Gifford Wallingford Oxfordshire OX10 8BB UK
3. Butterfly Conservation; Manor Yard East Lulworth Dorset BH20 5QP UK
4. Department of Crop and Environment Sciences; Harper Adams University College; Edgmond Newport Shropshire TF10 8NB UK
5. NERC Centre for Ecology and Hydrology; Bush Estate Penicuik Edinburgh EH26 0QB UK