1. School of Mathematics and StatisticsUniversity of Glasgow Glasgow UK
2. FitzPatrick Institute of African OrnithologyDST‐NRF Centre of ExcellenceUniversity of Cape Town Rondebosch South Africa
3. Integrated Research CenterThe Field Museum of Natural History Chicago Illinois
4. Tanzania Wildlife Research Institute Arusha Tanzania
5. Institute of Biodiversity, Animal Health and Comparative MedicineUniversity of Glasgow Glasgow UK