1. School of Mathematics and Statistics and Evolution & Ecology Research Centre; The University of New South Wales; Sydney NSW 2052 Australia
2. Centre for Behavioural and Physiological Ecology; Discipline of Zoology; School of Environmental and Rural Science; University of New England; Armidale NSW 2351 Australia
3. Evolution & Ecology Research Centre and School of Biological, Earth and Environmental Sciences; The University of New South Wales; Sydney NSW 2052 Australia
4. Department of Zoology; La Trobe University; Melbourne Vic. 3068 Australia