1. Colorado Cooperative Fish and Wildlife Research Unit, Department of Fish, Wildlife, and Conservation BiologyColorado State University Fort Collins CO USA
2. Department of StatisticsColorado State University Fort Collins CO USA
3. U.S. Geological Survey, Colorado Cooperative Fish and Wildlife Research Unit, Department of Fish, Wildlife, and Conservation BiologyColorado State University Fort Collins CO USA
4. National Park ServiceSoutheast Alaska Inventory and Monitoring Network Juneau AK USA
5. National Park ServiceGlacier Bay Field Station Juneau AK USA