1. Department of Mathematics and Statistics Lancaster University Lancaster UK
2. School of Mathematics and Maxwell Institute for Mathematical Sciences University of Edinburgh Edinburgh UK
3. Geography, Earth & Environmental Sciences University of Birmingham Birmingham UK
4. Biodiversity, Ecology & Conservation Group International Institute for Applied Systems Analysis Vienna Austria
5. Department of Biosciences Swansea University Swansea UK
6. Centre for Biomathematics Swansea University Swansea UK