1. Wyoming Cooperative Fish and Wildlife Research Unit, Department of Zoology and Physiology University of Wyoming Laramie Wyoming USA
2. Biology Department University of Washington Seattle WA USA
3. Idaho Department of Fish and Game Boise Idaho USA
4. U.S. Geological Survey, Northern Rocky Mountain Science Center West Glacier Montana USA
5. Program in Ecology University of Wyoming Laramie Wyoming USA
6. Western EcoSystems Technology (WEST), Inc. Laramie Wyoming USA
7. School of Life Sciences University of KwaZulu‐Natal Durban South Africa
8. Department of Zoology and Physiology University of Wyoming Laramie Wyoming USA
9. U.S. Geological Survey, Wyoming Cooperative Fish and Wildlife Research Unit, Zoology and Physiology Department University of Wyoming Laramie Wyoming USA