1. The Institute of Statistical Mathematics 10‐3 Midoricho, Tachikawa Tokyo 190‐8562 Japan
2. USGS Patuxent Wildlife Research Center 12100 Beech Forest Road Laurel MD 20708 USA
3. National Research Institute of Far Seas Fisheries Japan Fisheries Research and Education Agency 2‐12‐4 Fukuura, Kanazawa‐ku Yokohama Kanagawa 236‐8648Japan
4. Akkeshi Marine Station, Field Science Center for Northern Biosphere Hokkaido University Aikappu, Akkeshi Hokkaido 088‐1113 Japan
5. Faculty of Environmental Earth Science Hokkaido University N10W5, Kita‐ku Sapporo Hokkaido 060‐0810 Japan