1. Australian Research Council Centre of Excellence for Coral Reef Studies; James Cook University; Townsville QLD 4811 Australia
2. Chico Mendes Institute for Biodiversity Conservation; Ministry of Environment; Brasilia DF 70.670-350 Brazil
3. School of Environmental Studies; University of Victoria; Victoria BC V8W 2Y2 Canada
4. PSL Research University, CEFE UMR 5175, CNRS, Université de Montpellier; Université Paul-Valéry Montpellier, EPHE, Biogeographie et Ecologie des Vertébrés; Montpellier France
5. MARBEC UMR 9190, CNRS - IRD; Universite Montpellier - Ifremer; 34095 Montpellier France
6. Complex Systems Group, Department of Energy and Environment; Chalmers University of Technology; SE-412 96 Gothenburg Sweden