1. Graduate School of Environmental Science; Hokkaido University; Sapporo; 060-0810; Japan
2. Takehara Marine Science Station; Setouchi Field Science Centre, Graduate School of Biosphere Science, Hiroshima University, 5-8-1 Minato-mach; Takehara; Hiroshima; 725-0024; Japan
3. Department of Life Sciences; Natural History Museum, Cromwell Road; London; SW7 5BD; UK
4. Faculty of Science; Department of Geosciences, Shizuoka University, 836 Ohya; Shizuoka; 422-8529; Japan