1. Ornithological Section; Senckenberg Research Institute and Natural History Museum Frankfurt; Senckenberganlage 25 Frankfurt am Main D-60325 Germany
2. Vertebrate Palaeontology Laboratory; Department of Earth Sciences; University of Hong Kong; Pokfulam Hong Kong China
3. School of Biological Sciences; University of Bristol; Life Sciences Building 24 Tyndall Avenue Bristol BS8 1TQ UK
4. Burke Museum of Natural History and Culture; Seattle WA USA