1. Department of Biology; Duke University; Durham NC 27708 USA
2. Soil and Water Sciences Department; North Florida Research and Education Center; University of Florida; Quincy FL 32351 USA
3. School of Plant Sciences and Department of Ecology and Evolutionary Biology; University of Arizona; Tucson AZ 85721 USA
4. Department of Plant, Soil and Microbial Sciences; Michigan State University; East Lansing MI 48824 USA