1. State Key Laboratory of Marine Environmental Sciences Xiamen University Xiang'an Xiamen 361101 China
2. Institute of Marine Microbes and Ecospheres Xiamen University Xiang'an Xiamen 361101 China
3. Department of Limnology and Oceanography University of Vienna Althanstrasse 14 A‐1090 Vienna Austria
4. Department of Biological Oceanography Royal Netherlands Institute for Sea Research 1790 AB Den Burg Texel the Netherlands