1. Laboratory of Vaccine Materials, Center for Vaccine and Adjuvant Research and Laboratory of Gut Environmental System National Institutes of Biomedical Innovation, Health and Nutrition (NIBIOHN) Ibaraki‐city, Osaka Japan
2. Department of Microbiology and Immunology Kobe University Graduate School of Medicine Kobe‐city, Hyogo Japan
3. Graduate School of Pharmaceutical Sciences Osaka University Suita‐city, Osaka Japan
4. Innovation Center Nippon Flour Mills Co., Ltd Atsugi-city, Kanagawa Japan
5. Department of Dermatology Kyoto University Graduate School of Medicine Kyoto-city, Kyoto Japan
6. Laboratory of Proteome Research NIBIOHN Ibaraki‐city, Osaka Japan
7. International Research and Development Center for Mucosal Vaccines The Institute of Medical Science, The University of Tokyo Minato-ku, Tokyo Japan
8. Department of Immunology, Graduate School of Medicine Chiba University Chiba‐city, Chiba Japan
9. Division of Gastroenterology, Department of Medicine University of California, San Diego (UCSD) San Diego California
10. CU‐UCSD Center for Mucosal Immunology, Allergy and Vaccines (cMAV) UCSD San Diego California
11. Graduate School of Medicine and Graduate School of Dentistry Osaka University Suita‐city, Osaka Japan