1. School of Biological Sciences; University of Bristol; Life Sciences Building 24 Tyndall Avenue Bristol BS8 1TQ UK
2. UMR 1332 Biologie du Fruit et Pathologie; INRA; CS 20032 33882 Villenave d'Ornon Cedex France
3. UMR 1332 Biologie du Fruit et Pathologie; Université de Bordeaux; CS 20032 33882 Villenave d'Ornon Cedex France
4. The Food and Environment Research Agency; Sand Hutton York YO41 1LZ UK