1. Fisheries College; Ocean University of China; Qingdao Shandong China
2. The Key Laboratory of South China Sea Fishery Resources Exploitation and Utilization; Ministry of Agriculture; South China Sea Fisheries Research Institute; Chinese Academy of Fishery Sciences; Guangzhou Guangdong China
3. Engineering Research Center of Shellfish Culture and Breeding of Liaoning Province; College of Fisheries and Life Science; Dalian Ocean University; Dalian Liaoning China