1. Department of Molecular and Cell Biology; University of Cape Town; Rondebosch 7701 South Africa
2. School of Life Sciences; University of Warwick; Coventry CV4 7AL UK
3. Warwick Systems Biology Centre; University of Warwick; Coventry CV4 7AL UK
4. College of Life and Environmental Sciences; University of Exeter; Exeter EX4 4QD UK