1. GEODE UMR 5602 CNRS Université Jean‐Jaurès 5 Allée Antonio‐Machado Toulouse 31058 France
2. German Centre for Integrative Biodiversity Research Halle‐Jena‐Leipzig Deutscher Platz 5e Leipzig 04103 Germany
3. Centre for Biodiversity and Conservation Science School of Biological Sciences The University of Queensland Brisbane QLD 4072 Australia