1. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
2. Yunnan International Joint Laboratory for Biodiversity of Central Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
3. Department of Ecology and Evolutionary Biology University of Toronto 25 Willcocks Street Toronto ON M5S 3B2 Canada
4. Evolution & Ecology Research Centre, School of Biological, Earth and Environmental Sciences UNSW Sydney Sydney NSW 2052 Australia
5. School of Biological Sciences University of Portsmouth Portsmouth PO1 2DY UK
6. Institute of Arctic Biology University of Alaska Fairbanks Fairbanks AK 99775 USA
7. Yunnan Forestry Technological College Kunming 650224 China
8. State Key Laboratory of Urban and Regional Ecology, Research Center for Eco‐Environmental Sciences Chinese Academy of Sciences Beijing 100085 China
9. Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology, Institute of Biodiversity, School of Ecology and Environmental Science Yunnan University Kunming 650504 China