1. Department of Horticulture, Key Laboratory of Horticultural Plant Growth, Development and Quality improvement, Ministry of Agriculture Zhejiang University Hangzhou 310058 China
2. School of Horticulture and Landscape Architecture Yangzhou University Yangzhou 221116 China
3. Key Laboratory of Biotechnology in Plant Protection of Ministry of Agriculture and Zhejiang Province Zhejiang Academy of Agricultural Sciences Hangzhou 310021 China
4. College of Horticulture Sichuan Agricultural University Chengdu 611130 China