1. Department of Anatomy and Cell Biology; Medical Research Center Oulu; University of Oulu; Oulu Finland
2. Department of Archaeology; University of Oulu; Oulu Finland
3. Center for Functional Anatomy and Evolution; Johns Hopkins University School of Medicine; Baltimore MD USA
4. Centre for Life Course Epidemiology and Systems Medicine; Medical Research Center; University of Oulu and Oulu University Hospital; Oulu Finland
5. Finnish Institute of Occupational Health, Work and Health Ability and Disability Prevention Centre; Oulu Finland
6. Department of Diagnostic Radiology; Oulu University Hospital; Oulu Finland
7. Center for Medical Imaging; Physics and Technology Research; University of Oulu; Oulu Finland
8. Institute of Health Sciences, Public Health and General Practice; University of Oulu; Oulu Finland
9. Biocenter Oulu; University of Oulu; Oulu Finland
10. National Institute of Health and Welfare; Oulu Finland
11. Department of Biostatistics and Epidemiology; Faculty of Medicine; School of Public Health; Imperial College; London UK