1. Dept of Ecology, Environment and Plant Sciences, Stockholm Univ. SE‐10691 Stockholm Sweden
2. Environmental and Marine Biology, Åbo Akademi Univ. FI‐20520 Turku Finland
3. Dept of Aquatic Resources, Inst. of Coastal Research, The Swedish Univ. of Agricultural Sciences Öregrund Sweden
4. Åland Univ. of Applied Sciences, Mariehamn Åland Finland