1. Joint Faculty of Veterinary Medicine Department of Pathogenetic and Preventive Veterinary Science, Kagoshima University Kagoshima Japan
2. Kagoshima Crane Conservation Committee Izumi Kagoshima Japan
3. Joint Faculty of Veterinary Medicine Transboundary Animal Diseases Research Center, Kagoshima University Kagoshima Japan
4. Joint Graduate School of Veterinary Science Kagoshima University Kagoshima Japan
5. Matsuoka Research Institute for Science Tokyo Japan